An additional species of the genus Nemaspela Silhavý, 1966 from a cave in Georgia, with a note on Nemaspela melouri (Opiliones: Nemastomatidae).

A cavernicolous harvestman species, Nemaspela kotia sp. nov., is described from the Kotia Cave (=Kotiasklde Cave) in Zemo Imereti plateau, Imereti Region, West Georgia. Its males lack a cheliceral apophysis, a feature shared with the Georgian cave-dwelling Nemaspela femorecurvata Martens, 2006 and N. prometheus Martens, Maghradze & Barjadze, 2021. Besides, Nemaspela melouri Martens, Maghradze & Barjadze, 2021, known only from its type locality, is found in Solkota Cave (Tskaltubo Municipality, Sataplia-Tskaltubo karst massif, Imereti region, West Georgia). A key for the identification of the known species of Nemaspela is provided.

Diversity, taxonomy and phylogenetic relationships of the Niphargus borutzkyi ingroup (Crustacea: Amphipoda: Niphargidae) in Western Georgia, SW Caucasus.

Two new Caucasian species of the genus Niphargus Schidte, 1849 (Crustacea: Amphipoda: Niphargidae), N. rachalechkhumensis sp. nov. and N. tvishiensis sp. nov., are described from the stygobiotic habitats of the Racha-Lechkhumi and Kvemo Svaneti karst systems in Western Georgia. Both newly described species belong to the southwestern Caucasian Niphargus borutzkyi ingroup related to the European carphaticus species complex and can be clearly separated from the congeners by morphological features, mostly in uropod III and epimeral plates, and genetically. Identification key for all known species to the Niphargus borutzkyi ingroup is provided, as well as their phylogenetic relationships, the estimated time of the origin and the current distribution of the ingroup in the Colchis lowland of the southwestern Caucasus are also discussed.

Two new species of Plutomurus Yosii (Collembola: Tomoceridae) from the Caucasus and central Europe.

Plutomurus kharagauliensis sp. nov. from Kozmani Cave, Kharagauli Municipality, Georgia, and P. ruseki sp. nov. from the surface and subterranean habitats in different regions of Slovakia are described, illustrated and differentiated from other morphologically similar congeners. Both species are unique by dorsal macrochaetotaxy of head and abdomen. An identification key to the species of the genus Plutomurus reported from the Western Palearctic is presented.

The first troglobiotic cryptopid centipede (Chilopoda: Scolopendromorpha: Cryptopidae) from the Caucasus.

Cryptops (Cryptops) datviensis sp. nov. from Datvi Cave in Georgia, is the first troglobiotic species of the genus Cryptops Leach, 1814 to be described from the Caucasus. The new species morphologically resembles the common epigean Caucasian species Cryptops caucasius Verhoeff, 1934, but differs in typical troglomorphic features, such as elongation of antennae and legs. A key to species of the genus Cryptops from the Caucasus is presented. A list of the invertebrates inhabiting Datvi Cave is provided.

Two new species of the genus emNemaspela/em Silhavý from caves in Georgia (Opiliones: Nemastomatidae).

Two highly specialized endemic troglobiotic harvestman species of the genus Nemaspela Silhavý, 1966 are described. N. melouri sp. nov. from Melouri Cave and N. prometheus sp. nov. from Prometheus Cave (Sataplia-Tskaltubo karst massif, Imereti region, western Georgia), respectively. Despite the fact that the entrances of the caves are positioned only 2.5 km apart, the new taxa differ from each other distinctly by presence vs. absence of male cheliceral apophysis, which is lacking in the second species. A key to the Caucasian species of the genus is provided. Relationships of Nemaspela species within the genus and with hypothetical epigean ancestors are discussed.